Protochirotherium

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phylum Chordata Protochirotherium (‘first hand-beast’) is a Late Permian?-Early Triassic ichnotaxon (trace fossil) consisting of five-fingered footprints and whole tracks, discovered in Germany and later Morocco, Poland and possibly also Italy. The tracks were made by early archosauriforms with an erect posture. Erythrosuchids have been suggested as responsible for some European tracks, though they may have had a broader gait. Early pseudosuchian archosaurs are a more widely accepted candidate.

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clade Diapsida
Ichnogenus Protochirotherium
Temporal range Early Triassic